I came across another reference as to the function of gill rakers:
Trout, especially in lakes, also feed on zooplankton as small as 1 millimeter in diameter. Rather than take them in through the mouth, trout funnel them through their gill rakers, bony processes on the gill arches, into the esophogus.
This was taken from Trout, Stackpole Books. Jim Chacko Athappilly is a proffessor at Untity College in Maine, certified fish pathologist, etc. I should mention that he does list Behnke as a scource concerning the number of gill rakers in Kokanee!
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